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{{Unreferenced|date=July 2010}}
{{Weasel|date=July 2010}}
<!--http://www.europuppyusa.com/images/dog_breed_info/127_1.jpg
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{{Infobox Dogbreed
{{Infobox Dogbreed
| name = Tibetan Mastiff
| akcgroup = Working (January 2007)
| image = File:BIR Grupp 2- TIBETANSK MASTIFF, Legenda Tibeta Temudzhin (23938629470).jpg
| akcstd = http://www.akc.org/breeds/tibetan_mastiff/index.cfm
| image_caption =
| altname = Do-Khyi<br>Tsang-khyi<br>藏獒(Zang Ao)
| origin = Tibetan Plateau
| ankcgroup = Group 6 (Utility)
| maleheight = {{convert|66|cm|in|abbr=on|0}}<ref name=fci>[http://www.fci.be/Nomenclature/Standards/230g02-en.pdf FCI breed standard]</ref>
| ankcstd = http://www.ankc.aust.com/tibmast.html
| femaleheight = {{convert|61|cm|in|abbr=on|0}}<ref name=fci/>
| ckcgroup = Miscellaneous
| fcistd = http://www.fci.be/Nomenclature/Standards/230g02-en.pdf
| ckcstd = ?
}}
|ckcmisc=yes
The '''Tibetan Mastiff'''{{efn|{{lang-bo|འབྲོག་ཁྱི། འདོགས་ཁྱི། སེང་ཁྱི། |lit=Nomad-dog; chaining dog; lion-dog|translit=Drok chi/kyi; 'Dok kyi; Seng kyi|label=Tibetan}}, [[THL Simplified Phonetic Transcription|THL]]: '''{{transliteration|bo|Do khyi}}''',<!-- this is the correct spelling; do not change --> [[Wylie transliteration|Wylie]]: 'brog khyi; {{transliteration|bo|'dogs khyi}}; seng khyi}} is a large [[Tibetan dog breeds|Tibetan dog breed]]. Its double coat is medium to long, subject to climate, and found in a wide variety of colors, including solid black, black and tan, various shades of red (from pale gold to deep red) and bluish-gray (dilute black), and sometimes with white markings around its neck, chest and legs.
| country = [[China]]
| patronage = [[Fédération Cynologique Internationale|F.C.I.]] <ref name="FCI">[http://www.fci.be/uploaded_files/230A2004_en.doc Fédération Cynologique Internationale Standard] No. 230 of March 23, 2004, translated April 2, 2004, retrieved 2009-04-12 {{en icon}}</ref>
| fcigroup = 2
| fcinum = 230
| fcisection = 2
| fcistd = http://www.dogdomain.com/FCI/fcistandards/fci-230.htm
| image =Bea Miu Nan Šan, CAC.jpg
| image_caption =
| kcukgroup = Working
| kcukstd = http://www.thekennelclub.org.uk/item/63
| name = Tibetan Mastiff
| nzkcgroup = Utility
| nzkcstd = http://www.nzkc.org.nz/br697.html
| ukcgroup = Guardian Dogs
| ukcstd = http://mail.ukcdogs.com/UKCweb.nsf/80de88211ee3f2dc8525703f004ccb1e/de5952c978cfb24c85257044006b7b2c?OpenDocument
}}<!-- End Infobox Dogbreed info. Article Begins Here -->


==Name==
The '''Tibetan Mastiff''' (Do-khyi) is an ancient [[dog breed|breed]] and [[Dog type|type]] of [[domestic dog]] (''Canis lupus familiaris'') originating with nomadic cultures of [[Central Asia]].
[[File:Ten Prized Dogs 03.jpg|thumb|Ten Prized Dogs series, an artwork depicting a Tibetan mastiff.]]


The term ''[[mastiff]]'' was assigned by the Europeans who first came to Tibet because that name was used to refer to nearly all large dog breeds in the West. Early Western visitors to Tibet misnamed several of its breeds, such as the [[Tibetan Terrier|Tibetan terrier]], which is not a [[terrier]], and the [[Tibetan Spaniel|Tibetan spaniel]], which is not a [[spaniel]]. A better name for the breed might be the ''Tibetan mountain dog'' or—to encompass the landrace breed throughout its range—the ''Himalayan mountain dog''.<ref name=Messerchmidt/>
==Names and etymology==
==Description==
The Tibetan Mastiff also known as Do-khyi (variously translated as "home guard", "door guard", "dog which may be tied", "dog which may be kept"), reflects its use as a guardian of herds, flocks, [[tent]]s, villages, [[monasteries]], and [[palace]]s, much as the old English ban-dog (also meaning tied dog) was a dog tied outside the home as a guardian. However, in nomad camps and in villages, the Do-khyi is traditionally allowed to run loose at night and woe be unto the stranger who walks abroad after dark.
{{More citations needed section|date=February 2021}}

'Bhote Kukur' in [[Nepali language|Nepali]] means Tibetan Dog. In [[Mandarin Chinese]], the name is 藏獒 (Zang'Ao), which literally means Tibetan Mastiff or Tibetan "big ferocious dog". In [[Mongolia]] it is called "bankhar", meaning "guard dog". But there is an another type of Mastiff in Mongolia, called Mongolian Mastiff (Mongol Bankhar) which is bigger than Tibetian Mastiff and have darker color. The [[molosser]] type with which the modern Tibetan Mastiff breed is purportedly linked was known across the Ancient world by many names.

== Description ==


===Appearance===
===Appearance===
[[File:LegendaryTubo.jpg|thumb|This side profile of Tü-Bo underscores the soundness and type that made him a sought-after stud dog.]]
[[File:Tibetanmastiffskull.jpg|thumb|Illustration of a Tibetan mastiff skull by [[Frédéric Cuvier]]]]
It typically retains the hardiness which would be required for it to survive in Tibet, Ladakh and other high-altitude Himalayan regions.
Currently, some breeders differentiate between two "types" of Tibetan Mastiff: The Do-khyi and the "Tsang-khyi". The "Tsang-khyi" (which, to a Tibetan, means only "dog from Tsang") is also referred to as the "monastery type", described as generally taller, heavier, more heavily boned, with more facial wrinkling and haw than the "Do-khyi" or "nomad type". Both "types" are often produced in the same litter.


Instinctive behaviors, including canine pack behavior, contributed to the survival of the breed in harsh environments. It is one of the few primitive dog breeds that retains a single [[estrus]] per year instead of two, even at much lower altitudes and in much more temperate climates than its native climate. This characteristic is also found in wild canids such as the [[wolf]] and other wild animals. Since its estrus usually takes place during late autumn, most Tibetan Mastiff puppies are born between December and January.<ref>{{cite web |url=https://www.akc.org/dog-breeds/tibetan-mastiff/ |title=Tibetan mastiff |series=Dog breed information |website=American Kennel Club |access-date=9 March 2019}}</ref>
Males can reach heights up to 31+ inches (80+cm) at the [[withers]], although the standard for the breed is typically in the 25 to 28&nbsp;inch (61 to 72&nbsp;cm) range. The heaviest TM on record may be one weighing over 130&nbsp;kg (286.6 Lbs) {{Citation needed|date=February 2007}} but dogs bred in the West are more typically between 140&nbsp;lb (64&nbsp;kg) to 180&nbsp;lb (82&nbsp;kg)—especially if they are in good condition and not overweight. The enormous dogs being produced in some Western and some Chinese kennels would have "cost" too much to keep fed to have been useful to nomads; and their questionable structure would have made them well-nigh useless as livestock guardians.


Its double coat is long, subject to climate, and found in a wide variety of colors, including solid black, black and tan, various shades of red (from pale gold to deep red) and bluish-gray (dilute black), often with white markings. {{As of|2014}}, some breeders had begun marketing white Tibetan mastiffs.
The Tibetan Mastiff is considered a primitive breed. It typically retains the instincts which would be required for it to survive in Tibet, including canine pack behaviour. In addition, it is one of the few primitive dog breeds that retains a single oestrus per year instead of two, even at much lower altitudes and in much more temperate climates than its native climate. This characteristic is also found in wild canids such as the [[wolf]]. Since its [[oestrus]] usually takes place during late fall, most Tibetan Mastiff puppies are born between December and January.<ref>[http://www.akc.org/breeds/tibetan_mastiff/did_you_know.cfm American Kennel Club - Tibetan Mastiff Did You Know?<!-- Bot generated title -->]</ref>


The coat of a Tibetan Mastiff lacks the unpleasant big-dog smell that affects many large breeds. The coat, whatever its length or color(s), should shed dirt and odors. Although the dogs shed somewhat throughout the year, there is generally one great molt in late winter or early spring and sometimes another, lesser molt in the late summer or early autumn. (Sterilization of the dog may dramatically affect the coat as to texture, density and shedding pattern.)<ref>citation needed</ref>
[[File:Mastif tybetański 2009 pl3.jpg|thumb|Tibetan Mastiff at an international dog show in Poland.]]
Its double coat is long, subject to climate, and found in a wide variety of colors, including solid black, black & tan, various shades of gold, blue/gray, chocolate brown, the rarest being solid white.


Tibetan Mastiffs are shown under one standard in the West, but separated by the Indian breed standard into two varieties: Lion Head (smaller; exceptionally long hair from forehead to withers, creating a ruff or mane) and Tiger Head (larger; shorter hair).
The coat of a Tibetan Mastiff lacks the unpleasant "big-dog smell" that affects many large breeds. The coat, whatever its length or color(s), should shed dirt and odors. Although the dogs shed somewhat throughout the year, there is generally one great "molt" in late winter or early spring and sometimes another, lesser molt in the late summer or early fall. (Sterilization of the dog or bitch may dramatically affect the coat as to texture, density, and shedding pattern.)

Tibetan Mastiffs are shown under one standard in the West, but separated by the Indian breed standard into two varieties:{{Citation needed|date=February 2007}} Lion Head (smaller; exceptionally long hair from forehead to withers, creating a ruff or mane) and Tiger Head (larger; shorter hair).


===Temperament===
===Temperament===
As a [[Livestock guardian dog|flock guardian dog]] in Tibet, and in the West, it uses all the usual livestock guardian tactics (e.g., barking, scent-marking perimeters) to warn away predators and avoid direct confrontations.<ref name=Messerchmidt/>


As a socialized, more domestic dog, it can thrive in a spacious, fenced yard with a canine companion, but it is not an appropriate dog for apartment living. The Western-bred dogs are generally more easy-going, although still somewhat aloof with strangers. Through hundreds of years of selective breeding for a protective flock and family guardian, the breed has been prized for being a nocturnal sentry, keeping would-be predators and intruders at bay, and barking at unidentified sounds throughout the night. Leaving a Tibetan Mastiff outside all night with neighbors nearby is not recommended. They often sleep during the day, making them more active, alert, and aware at night.<ref name=Messerchmidt/>
The native type of dog, which still exists in [[Tibet]], and the Westernized purebred breed can vary in temperament—but so can dogs of identical breeding, within the same litter, raised in the same household. Elizabeth Schuler states, "The few individuals that remain in Tibet are ferocious and aggressive, unpredictable in their behavior, and very difficult to train. But the dogs bred by the English are obedient and attached to their masters." However, other observers have found the dogs remaining in Tibet to be quite approachable under the right circumstances—and some Western-bred dogs to be completely unapproachable.


Like all flock guardian breeds, they are intelligent and stubborn to a fault, so [[obedience training]] is recommended (although it is only mildly successful with some individuals) since this is a strong-willed, powerful-bodied breed. Unless they are to be used exclusively as livestock guardians, [[Socialization of animals|socialization]] training is also critical with this breed, because of their reserved nature with strangers and guardian instincts. They can be excellent family dogs, depending on the family; owners must understand canine psychology and be able and willing to spend a lot of time and devotion to training their dogs. Lack of consistency can result in the creation of unpredictable dogs. The protectiveness of Tibetan Mastiffs requires alertness and planning by the owner, in order to avoid mishaps when the dog is merely reacting as a guardian. The breed is not recommended for novice dog owners.<ref name=Messerchmidt/><ref name=Anthology/>
Some Western and Asian breeders are seeking to create a replica of the legendary dog which they identify as the "true Tibetan Mastiff" or "Tsang-khyi". Some breeders appear to select primarily for appearance (great size, profuse coat, heavy wrinkling, jowls, haw) while others also select for "soft" temperament (in the West) and fierce temperament (in Asia where the dogs' "ferocity" is much vaunted and encouraged).

As a [[Livestock guardian dog|flock guardian dog]] in Tibet and in the West, it is tenacious in its ability to confront predators the size of [[wolf|wolves]] and [[leopard]]s. As a socialized, more domestic dog, it can thrive in a spacious, fenced yard with a canine companion, but it is generally not an appropriate dog for apartment living. The Western-bred dogs are generally more easy-going, although somewhat aloof with strangers coming to the home. Through hundreds of years of selective breeding for a protective flock and family guardian, the breed has been prized for being a nocturnal sentry, keeping would-be predators and intruders at bay, barking at sounds throughout the night. Leaving a Tibetan Mastiff outside all night with neighbors nearby is not recommended. They often sleep during the day to be more active, alert and aware at night.

Like all flock guardian breeds, they are intelligent and stubborn to a fault, so [[obedience training]] is recommended (although only mildly successful with some individuals) since this is a strong-willed, powerful breed. [[Socialization]] is also critical with this breed because of their reserved nature with strangers and guardian instincts. They are excellent family dogs—for the right family. Owners must understand canine psychology and be willing and able to assume the primary leadership position. Lack of consistent, rational discipline can result in the creation of dangerous, unpredictable dogs(although this is true of virtually every dog breed).

Newspaper reports have suggested that a pair of these Mastiffs have killed tigers while guarding sheep in the highlands of Nepal.


==Health==
==Health==
{{More citations needed section|date=February 2021}}
[[File:Zangao.jpg|thumb|A Chinese bred Tibetan Mastiff]]


'''Life Expectancy''' Unlike most large breeds, its [[life expectancy]] is long, some 10–14 years. The breed has fewer genetic health problems than many breeds, but cases can be found of [[hypothyroidism]], [[entropion]], [[ectropion]], skin problems including [[allergy|allergies]], autoimmune problems including [[demodex]], missing teeth, [[malocclusion]] ([[overbite]] or [[underbite]]), cardiac problems, [[epilepsy]], [[progressive retinal atrophy]] (PRA), [[cataract]], and small ear canals with a tendency for infection. As with most large breeds, some will suffer with elbow or [[Hip dysplasia (canine)|hip dysplasia]], although this has not been a major problem in the Tibetan Mastiff. Another concern includes [[Polyneuropathy in dogs and cats|canine inherited demyelinative neuropathy]] (CIDN), a rare inherited [[neural]] disease that appeared in one bloodline in the early 1980s.
[[life expectancy]] long, . The breed has fewer genetic health problems than many breeds, but cases can be found of [[hypothyroidism]], [[entropion]], [[ectropion]], skin problems including [[|allergies]], autoimmune problems including [[demodex]], missing teeth, [[malocclusion]] ([[overbite]] [[underbite]]), cardiac problems, [[epilepsy]], [[progressive retinal atrophy]] (PRA), [[cataract]], and small ear canals with a tendency for infection. As with most large breeds, some will suffer with elbow or [[Hip dysplasia (canine)|hip dysplasia]].


Canine inherited demyelinative neuropathy (CIDN), an inherited condition, appeared in one of the prominent lines of Tibetan Mastiffs in the early 1980s.<ref>"The Tibetan Mastiff" by Ann Rohrer and Cathy J. Flamholtz</ref>{{full citation needed|date=January 2021}} Unfortunately, known carriers were bred extensively and are behind many lines still being actively bred.{{cn|date=September 2022}} Because the mode of inheritance appears to be as a simple recessive, continued inbreeding can still produce affected puppies.
'''Canine Hip Dysplasia''' (CHD) takes many forms, e.g., the femoral head ("ball") may not fit well into the acetabulum ("socket"); the ligament connecting the two may be lax, allowing dislocation; there may be no femoral head at all. Not all forms cause clinical signs. Very active, well-muscled dogs with no femoral heads may show no impairment. Their owners may be unaware of their dogs' "hip dysplasia" unless/until there is a reason to x-ray the hips.


Hypothyroidism is fairly common in Tibetan Mastiffs, as it is in many large "northern" breeds. They should be tested periodically throughout their lives using a complete thyroid "panel". However, because the standard thyroid levels were established using domestic dog breeds, test results must be considered in the context of what is "normal" for the breed, not what is normal across all breeds. Many dogs of this breed will have "low" thyroid values, but no clinical symptoms. Vets and owners differ on the relative merits of medicating dogs which test "low", but are completely asymptomatic.
As with all dog breeds, hip dysplasia is caused by the interaction of genes and environment. Inheritance of CHD appears to be polygenic, i.e., it is caused by more than one gene. Mode of inheritance (dominant, recessive, dominant with incomplete penetrance, etc.)has not been determined but may be different in different breeds. Rapid growth and weight gain in puppies may trigger or exacerbate a genetic tendency to all sorts of skeletal problems. Many TM breeders recommend against feeding "puppy food" and especially against feeding "large-breed" puppy food, as these concoctions may contain too many calories, leading to fat puppies. Some breeders and owners believe that supplementation with Vitamin C may prevent the development of CHD even in dogs with the genes for it.


==History==
'''Canine Inherited Demyelinative Neuropathy''' is an inherited condition that appeared in one of the prominent lines of Tibetan Mastiffs in the early 1980s. CIDN affect the peripheral nervous system. Nerve fibers are unable to transmit impulses from the spinal cord to the muscles because of the breakdown of the myelin sheath. Starting at approximately six weeks of age, affected pups begin to lose the ability to walk or even stand. Progression of the condition can take anywhere from a few days to two weeks.<ref>"The Tibetan Mastiff" by Ann Rohrer and Cathy J. Flamholtz</ref>
[[File:142. Thibet Dog.JPG|thumb|left|Tibetan dog from the 1850s]]
[[File:Tibetdog1904.jpg|thumb|Realistic representation/correction of Tibetan dog in 1904 "The German dogs". Heliogravure, Richard Strbel]]


Originally these dogs were used to protect Buddhist monasteries and monks of Tibet from animals such as bears, wolves and snow leopards.<ref name=Palika>{{cite book |last=Palika |first=Liz |year=2007 |title=The Howell Book of Dogs: The definitive reference to 300&nbsp;breeds and varieties |publisher=John Wiley & Sons |page=[https://archive.org/details/howellbookdogsde00pali/page/n388 374] |isbn=978-0-470-17585-9 |url=https://archive.org/details/howellbookdogsde00pali |url-access=registration |via=archive.org}}</ref><ref name=Messerchmidt>{{cite book |title=Discovering the Big Dogs of Tibet and the Himalayas: A personal journey |last=Messerchmidt |first=Don |year = 2010}} {{page needed |date=June 2020}}</ref><ref name=Anthology>{{cite book |title=Tibetan Dogs: A complete anthology of the breeds |date=18 November 2010 |publisher=Vintage Dog Books |isbn=978-1-4455-2671-3}} {{page needed|date=June 2020}}</ref>
Because this condition is inherited as a simple autosomal recessive, it is virtually impossible to completely eliminate it from the gene pool. One known carrier was bred to over 30 times, producing at least 134 direct descendants. Many descendants of this dog are still being bred so there is always the risk—however slim—of producing more affected puppies. Breeders need to be cautious about pairing up any two descendants of this dog.
'''Hypothyroidism''' is fairly common in Tibetan Mastiffs, as it is in many large "Northern" breeds. TMs should be tested periodically throughout their lives using a complete thyroid "panel". (Simple T2/T4 testing is virtually useless.) However, because the standard thyroid levels were established using domestic dog breeds, test results must be considered in the context of what is "normal" for the breed, not what is normal across all breeds. Many TMs will have "low" thyroid values but no clinical symptoms. Vets—and owners—differ on the relative merits of medicating dogs who "test low" but are completely asymptomatic. Some researchers think that asymptomatic hypothyroidism may have been adaptive in the regions of origin for many breeds, since less nutrition is required for the dog to stay in good condition. Therefore, attempts to eliminate "low thyroid" dogs from the TM gene pool may have unintended consequences for the breed.


The Tibetan Mastiff is a [[Phenotype|phenotypically]] distinct dog breed that was bred as a flock guardian in the high altitudes of the [[Himalayas]] and the [[Tibetan Plateau]]s.<ref>{{cite magazine |last=Messerschmidt |first=D.M.R. |year=1983 |title=The Tibetan Mastiff: Canine sentinels of the range |magazine=Rangelands |volume=5 |pages=172–174}}</ref><ref>{{cite journal |last1=Li |first1=Q. |last2=Liu |first2=Z. |last3=Li |first3=Y. |last4=Zhao |first4=X. |last5=Dong |first5=L. |last6=Pan |first6=Z. |last7=Sun |first7=Y. |last8=Li |first8=N. |last9=Xu |first9=Y. |last10=Xie |first10=Z. |year=2008 |title=Origin and phylogenetic analysis of Tibetan mastiff based on the mitochondrial DNA sequence |journal= Journal of Genetics and Genomics|volume=35 |issue=6 |pages=335–340 |doi=10.1016/S1673-8527(08)60049-1 |pmid=18571121 }}</ref>
In affected dogs, symptoms may include decreased activity and playing, increased sleeping, weight gain, poor skin and coat condition such as flaking and scaling, a "yeasty" smell to the coat, frequent ear infections, and negative changes in temperament. Fortunately, this condition is easily treated by the use of daily thyroid supplementation.


Meer Izzut-oollah (1872) wrote of the Tibetan Mastiff:
'''Osteochondritis Dessicans''' is a skeletal defect in which the cartilage lifts off the bone, becomes thickened and cracked, causes inflammation and pain, and in severe cases degeneration of the joint. This conditions strikes males more than females. Keeping an affected puppy lean may help but surgery may be required to relieve pain.


{{quote|The dogs of [[Tibet]] are twice the size of those seen in India, with large heads and hairy bodies. They are powerful animals...During the day they are kept chained up, and are let loose at night to guard their masters' house.<ref name=Izzut-oollah-1872>{{cite book |first=Meer |last=Izzut-oollah |year=1872 |title=Travels in Central Asia in the Years 1812–13 |translator=Henderson |publisher=Foreign Department Press |place=Calcutta, IN |page=15 |url=https://archive.org/details/in.gov.ignca.21243/mode/2up}}</ref>}}
'''Panosteitis''' is inflammation of the bones that strikes young dogs. The animal will become lame in one leg and then the inflammation will shift to a different leg. This is one condition that corrects itself over time, and only pain medication is needed.


In the early 20th century, the [[George V|Prince of Wales, the future George V]], introduced a pair of Tibetan Mastiffs to the United Kingdom. By 1906, the breed was prevalent enough in England to be shown at [[the Crystal Palace]] show. However, during the war years, the breed lost favor and focus and nearly died out in England.
'''Hypertrophic Osteodystrophy''' (HOD) is a condition that affects young large breed dogs. It is very painful and prognosis is fair to poor due to recurring episodes of the condition. Clinical signs of HOD include fever, lack of appetite, and depression. Lameness may vary from mild to severe. With multiple limbs affected, the dog may be reluctant to stand or walk. HOD may be mistaken for Panosteitis without proper diagnosis.


The breed has been gaining in popularity worldwide since 1980. Although the breed is still considered somewhat uncommon, as more active breeders arose and produced adequate numbers of dogs, various registries and show organizations (FCI, AKC) began to recognize the breed. In 2008, the Tibetan Mastiff competed for the first time in the [[Westminster Kennel Club Dog Show]].
Treatment is only supportive. Intravenous fluids are usually required to keep the patient hydrated. Nutritional support is provided with a feeding tube if the dog refuses to eat for five or more days. Pain is controlled with narcotics and NSAIDs. Antibiotics are used if the dog has signs of pneumonia or other bacterial infections. If the bones become twisted due to growth plate damage, corrective surgery may be indicated. Because the distemper vaccination has been implicated, inoculation should be delayed until the pet has been in remission for a couple of months. Information from http://www.vetsurgerycentral.com/hod.htm


Since AKC recognition, the number of active breeders has skyrocketed, leading to over-breeding of puppies, many of which are highly inbred and of questionable quality. Initially, the breed suffered because of the limited gene pool from the original stock.{{citation needed|date=January 2021}} By 2015, due to excessive breeding and unsuitability of the breed as a pet in urban situations, prices in China for the best dogs had fallen to about $2,000, and both lower quality and crossbreed dogs were being abandoned.<ref name="NYT41715">{{cite news |title=Once-prized Tibetan mastiffs are discarded as fad ends in China |date=2015-04-18 |series=World/Asia |newspaper=[[The New York Times]] |place=New York, NY |url=https://www.nytimes.com/2015/04/18/world/asia/once-prized-tibetan-mastiffs-are-discarded-as-fad-ends-in-china.html?_r=0 |access-date=27 February 2017}}</ref><ref name=Anthology/>
'''Ear Infections''' can be serious and the dog should be taken to the vet if you see it shaking its head or scratching more than normal. Tibetan Mastiffs have pendant ears, making them more prone to ear infections. The vet needs to determine the cause, and may prescribe antibiotics and/or ear drops. Some ear infections are contagious to other dogs if they involve mites or some bacteria.


In 2011, a DNA study concluded that there was a genetic relationship between the Tibetan mastiff and the [[Great Pyrenees]], [[Bernese Mountain Dog]], [[Rottweiler]] and [[St. Bernard (dog)|Saint Bernard]], and that these large breed dogs are probably partially descended from the Tibetan Mastiff.<ref>{{cite journal |last1=Li |first1=Y. |last2=Zhao |first2=X. |last3=Pan |first3=Z. |last4=Xie |first4=Z. |last5=Liu |first5=H. |last6=Xu |first6=Y. |last7=Li |first7=Q. |year=2011 |title=The origin of the Tibetan mastiff and species identification of ''Canis'' based on mitochondrial cytochrome&nbsp;c oxidase subunit&nbsp;I (COI) gene and COI barcoding |journal=Animal |volume=5 |issue=12 |pages=1868–73 |doi=10.1017/S1751731111001042 |doi-access=free |pmid=22440462|bibcode=2011Anim....5.1868L }}</ref> In 2014, a study added the [[Leonberger]] to the list of possible relatives.
==History==
[[File:142. Thibet Dog.JPG|thumb|Tibetan dog from the 1850s]]
This is an ancient breed. It has been theorized that an early Tibetan dog is the ancestor to all [[Molossus (dog)|Molossuses]] breeds. A study at Nanjing Agricultural University's Laboratory of Animal Reproductive Genetics and Molecular Evolution in Nanjing, China, found that while most common dog breeds genetically diverged from the wolf approximately 42,000 years ago, the Tibetan Mastiff genetically diverged from the wolf approximately 58,000 years ago.<ref>[http://www.sciencedirect.com/science?_ob=ArticleURL&_udi=B8H15-4SSRK5S-3&_user=10&_coverDate=06%2F30%2F2008&_rdoc=1&_fmt=high&_orig=search&_sort=d&_docanchor=&view=c&_searchStrId=1385511533&_rerunOrigin=google&_acct=C000050221&_version=1&_urlVersion=0&_userid=10&md5=77ecd1f1f64acdc5ef96bcc937fd3534 Origin and phylogenetic analysis of Tibetan Mastiff based on the mitochondrial DNA sequence<!-- Bot generated title -->]</ref> They share many characterisitcs of many [[Mountain dog]] breeds.


===Admixture with an unknown wolf-like canid===
Many Tibetan Mastiff breeders and owners (and their web sites) claim that [[Marco Polo]] encountered the large Tibetan dogs in his travels and described them as "tall as a donkey with a voice as powerful as that of a lion." However, reading of Polo's works does not support this. In fact, other travels told Marco Polo about these enormous dogs—and about unicorns and other exotic creatures.
The Tibetan Mastiff was able to adapt to the extreme highland conditions of the [[Tibetan Plateau]] very quickly compared with other mammals such as the [[Wild yak|yak]], [[Tibetan antelope]], [[snow leopard]], and the [[wild boar]]. The Tibetan mastiff's ability to avoid [[Hypoxic hypoxia|hypoxia]] in high altitudes, due to its higher [[hemoglobin]] levels compared with low-altitude dogs, was due to prehistoric interbreeding.<ref>{{cite journal |last1=Miao |first1=Benpeng |last2=Wang |first2=Zhen |last3=Li |first3=Yixue |year=2016 |title=Genomic analysis reveals hypoxia adaptation in the Tibetan mastiff by introgression of the grey wolf from the Tibetan plateau |journal=Molecular Biology and Evolution |volume=34 |issue=3 |pages=734–743 |s2cid=47507546 |doi=10.1093/molbev/msw274 |doi-access=free |pmid=27927792 |url=https://zenodo.org/record/895655}}</ref><ref>{{cite journal |last1=Signore |first1=Anthony V. |last2=Yang |first2=Ying-Zhong |last3=Yang |first3=Quan-Yu |last4=Qin |first4=Ga |last5=Moriyama |first5=Hideaki |last6=Ge |first6=Ri-Li |last7=Storz |first7=Jay F. |year=2019 |title=Adaptive changes in hemoglobin function in high-altitude Tibetan canids were derived via gene conversion and introgression |journal=Molecular Biology and Evolution |volume=36 |issue=10 |pages=2227–2237 |pmc=6759075 |doi=10.1093/molbev/msz097 |pmid=31362306}}</ref> In 2020, a genomic analysis indicates that a [[ghost population]] of an unknown wolf-like canid which is deeply-diverged from modern [[Wolf|Holarctic wolves]] and dogs has contributed the [[EPAS1]] allele found in both [[Himalayan wolf|Himalayan wolves]] and dogs, and this allows them to live in high altitudes.<ref>{{cite journal|doi=10.1093/molbev/msaa113|title=Ancient Hybridization with an Unknown Population Facilitated High-Altitude Adaptation of Canids|year=2020|last1=Wang|first1=Ming-Shan|last2=Wang|first2=Sheng|last3=Li|first3=Yan|last4=Jhala|first4=Yadvendradev|last5=Thakur|first5=Mukesh|last6=Otecko|first6=Newton O.|last7=Si|first7=Jing-Fang|last8=Chen|first8=Hong-Man|last9=Shapiro|first9=Beth|last10=Nielsen|first10=Rasmus|last11=Zhang|first11=Ya-Ping|last12=Wu|first12=Dong-Dong|journal=Molecular Biology and Evolution|volume=37|issue=9|pages=2616–2629|pmid=32384152|doi-access=free}}</ref>


==Gallery==
In the early 19th century, [[George IV of the United Kingdom|King George IV]] owned a pair of TMs, and there were enough of the breed in England in 1906 to be shown at the 1906 [[the Crystal Palace|Crystal Palace]] show. However, during the war years, the breed lost favor and focus and nearly died out in [[England]].
<gallery widths="170" heights="170">
File:Adventure, sport and travel on the Tibetan steppes (1911) (14597105527).jpg|Tibetan Mastiff with its owner in 1911
File:Tibetan Mastiff (2642055070).jpg|Tibetan Mastiff in Drepung Monastery, Lhasa, Tibet
File:Phinju.jpg|Tibetan Mastiff in Sandakpur, Nepal
File:Pyakurel.jpg|Tibetan Mastiff from Nepal
File:Tybetan Matt.jpg|Tibetan Mastiff
File:Tibetan Mastiff 001.jpg
File:Lion-Head Tibetan Mastif02.jpg|Tibetan Mastiff with Red Khekhor
File:Tibetan Mastiff.jpg|The Tibetan Mastiff is a livestock guard-dog
File:Tibet-5805 - Tibet at 15,000 feet (2589963541).jpg|Tibetan man with his Mastiff
</gallery>


==Popular culture==
After 1980, the breed began to gain in popularity worldwide. Although the breed is still considered somewhat uncommon, as various registries and show organizations (FCI, AKC) began to recognize the breed, more and more active breeders have arisen. Initially the breed suffered because of the limited gene pool from the original stock, but today's reputable breeders work hard at reducing the genetic problems through selective breeding and the international exchange of new bloodlines.
<!-- This is NOT a collection of any passing mentions of the breed; please only add significant referenced items! -->
{{wikinews|Four new breeds in the Westminster Kennel Club Dog Show.}}
In 2008, the Tibetan Mastiff competed for the first time in the [[Westminster Kennel Club Dog Show]].
Tibetan Mastiff the in the [[ ]].
* Ulysses, one of the main characters in the 2008 Christian animated film ''At Jesus' Side'', is a Tibetan Mastiff.
* A Tibetan Mastiff is the subject of the 2011 animated film ''[[The Tibetan Dog]]''.
*[[The Dresden Files characters#Mouse|Mouse]], a "Tibetan Temple dog" (a semi-divine creature that closely resembles a Mastiff), is the canine companion of the [[Harry Dresden|titular character]] of the ''[[The Dresden Files|Dresden Files]]'' book series.
* The animated film ''[[Rock Dog]]'' featured two Tibetan Mastiffs named "Bodi" and "Khampa" (voiced by [[Luke Wilson]] and [[J. K. Simmons]], respectively).
* In the 2018 animated television series, ''[[Craig of the Creek]]'', the character Wildernessa rides around on a Tibetan Mastiff named Cheesesticks.
* In the 2011 film ''[[Old Dog]]'' by director [[Pema Tseden]], the Tibetan Mastiff of a herder family is coveted by several characters to be traded to rich Han Chinese in the eastern part of the country.
* The 2019 video game [[Indivisible (video game)|''Indivisible'']] features Lanshi, a Tibetan Mastiff, as a playable character.
* The 2019 manga ''[[Weed (manga)#Ginga Densetsu Noah|Ginga Densetsu Noah]]'' features three Tibetan Mastiffs as the main antagonists.
* The 2020 video game ''[[Ghost of Tsushima]]'', Tibetan Mastiff is also known Bankhar Dog, were introduced as guard dog of Mongol army.
* The [[Pokémon]] character Mabosstiff, first featured in [[Pokémon Scarlet and Violet|Scarlet and Violet versions]], is heavily based on a Tibetan Mastiff.


==See also==
In recent years, more wealthy Chinese are buying Tibetan Mastiff as a means to show off their status. It was reported in Sep 2009<ref>{{cite web|url=http://www.wayangtimes.com/rich-chinese-woman-pays-more-than-us500000-for-rare-tibetan-mastiff-dog.html|title=Rich Chinese Woman Pays More than US$500,000 for Rare Tibetan Mastiff Dog}}</ref> that a rich Chinese woman spent more than USD 600,000 or 4 million yuan to buy an 18 month old purebred male Tibetan Mastiff, which she named Yangtze No. 2. This is—purportedly—the most ever paid for a dog. In December 2010, a buyer from Dalian paid 16 million yuan (almost US$2.5 million) for a slightly less than one-year-old puppy.
{{Portal|Dogs}}
* [[List of dog breeds]]
* [[Tibetan kyi apso|Tibetan Kyi Apso]]
* [[Dog of Osu]]
* [[Himalayan Sheepdog]]
* [[Lhasa Apso]]
* [[Tibetan Spaniel]]
* [[Tibetan Terrier]]
* [[Chow Chow]]


==Notes==
== Tibetan Mastiffs in popular culture ==
{{notelist}}
* A Tibetan Mastiff named Max is seen in the 1993 horror film, ''[[Man's Best Friend (1993 film)|Man's Best Friend]]''.
* In ''[[Blood Rites]]'', an novel of ''[[The Dresden Files]]'' by [[Jim Butcher]], the main character, [[Harry Dresden]], is "adopted" by a Tibetan mastiff. He names the dog "Mouse", who becomes a stalwart protector of the wizard.

== See also ==
* [[Dog]]
* [[Index of Tibet-related articles]]
* [[Dog of Osu]]


==References==
==References==
{{reflist}}
{{reflist}}
<!-- Please note the following are not Further reading. Please do not insert a heading for them. They are correctly listed after the citation list according to [[MOS:APPENDIX]] -->
* Alderton, David (1984). ''The Dog''.
* Fogle, Bruce, DVM (2000). ''The new Encyclopedia of the Dog''. Doring Kindersley (DK). ISBN 0-7894-6130-7.
* , (). ''The Dog''. . ISBN0---.
* Palmer, Joan (1994). ''The Illustrated Encyclopedia of Dog Breeds''. ISBN 0-7858-0030-1.
* , (). ''The Encyclopedia of ''. 0---.
* {{cite news |title= Tibetan mastiff, 'Big Splash', becomes world's most expensive dog |url=http://www.huffingtonpost.com/2011/03/16/tibetan-mastiff-most-expensive-dog-big-splash_n_836405.html |newspaper=[[The Huffington Post]] |date=May 25, 2011 |access-date=December 6, 2012}}
* Schuler, Elizabeth Meriwether (Ed.) (1980). ''Simon & Shuster's Guide to Dogs''. ISBN 0-671-25527-4.
* 倪正,(2000). "真实的藏獒 (The Real Tibetan Mastiff)". ISBN 7-80173-535-8
* 倪(2000). 真实的藏獒 The Tibetan . 7-80173-535-8
* Messerschmidt, Don (2010). ''Big Dogs of Tibet and the Himalayas''. Orchid Press {{ISBN|978-974-524-130-5}}.
* Palmer, Joan (1994). ''The Illustrated Encyclopedia of Dog Breeds''. {{ISBN|0-7858-0030-1}}.
* Schuler, Elizabeth Meriwether (ed.) (1980). ''Simon & Schuster's Guide to Dogs''. {{ISBN|0-671-25527-4}}.
* {{cite web |url=http://www.sify.com/news/world-s-most-expensive-dog-costs-a-million-pounds-news-international-ldqlkebbhci.html |archive-url=https://web.archive.org/web/20110319205135/http://www.sify.com/news/world-s-most-expensive-dog-costs-a-million-pounds-news-international-ldqlkebbhci.html |url-status=dead |archive-date=March 19, 2011 |title=World's most expensive dog costs a million pounds |date=March 16, 2011 |website=[[Sify]] |agency=[[Indo-Asian News Service|IANS]] |access-date=December 6, 2012}}


==External links==
==External links==
{{Commons category}}
{{Commonscat}}
{{Wikinews|Four new breeds in the Westminster Kennel Club Dog Show}}
* [http://www.huffingtonpost.com/2010/08/06/tibetan-mastiff-craze-swe_n_673763.html Tibetan Mastiff Craze Sweeps China]—slideshow by ''[[The Huffington Post]]''
<!--Editors: breed registry, club, rescue, and kennel information links are placed on DMOZ, not here-->
* {{curlie|Recreation/Pets/Dogs/Breeds/Livestock_Guardian_Dogs/Tibetan_Mastiff}}
* [http://www.huffingtonpost.com/2010/08/06/tibetan-mastiff-craze-swe_n_673763.html Tibetan mastiff craze sweeps China]—slideshow by ''[[The Huffington Post]]''


{{Pastoral dogs}}
{{ }}
{{Chinese dogs}}
{{Authority control}}


[[Category:Dog breeds originating in Tibet]]
[[Category:Dog breeds originating in Tibet]]
[[Category:Livestock guardians]]
[[Category: ]]
[[Category:Molossers]]
[[Category:]]
[[Category:Rare dog breeds]]

[[bo:དོ་ཁྱི།]]
[[bg:Тибетски дог]]
[[ca:Mastí tibetà]]
[[cs:Tibetská doga]]
[[da:Tibetansk mastiff]]
[[de:Do Khyi]]
[[es:Dogo del Tíbet]]
[[fr:Dogue du Tibet]]
[[it:Mastino tibetano]]
[[hu:Tibeti masztiff]]
[[nl:Tibetaanse mastiff]]
[[ne:भोटे कुकुर]]
[[ja:チベタン・マスティフ]]
[[no:Tibetansk mastiff]]
[[pl:Mastif tybetański]]
[[pt:Mastim tibetano]]
[[ru:Тибетский мастиф]]
[[fi:Tiibetinmastiffi]]
[[sv:Tibetansk mastiff]]
[[uk:Тибетський мастиф]]
[[vi:Chó ngao Tây Tạng]]
[[zh:藏獒]]

Latest revision as of 11:58, 29 September 2024

Tibetan Mastiff
OriginTibetan Plateau
Traits
Height Males 66 cm (26 in)[1]
Females 61 cm (24 in)[1]
Kennel club standards
Fédération Cynologique Internationale standard
Dog (domestic dog)

The Tibetan Mastiff[a] is a large Tibetan dog breed. Its double coat is medium to long, subject to climate, and found in a wide variety of colors, including solid black, black and tan, various shades of red (from pale gold to deep red) and bluish-gray (dilute black), and sometimes with white markings around its neck, chest and legs.

Name

[edit]
Ten Prized Dogs series, an artwork depicting a Tibetan mastiff.

The term mastiff was assigned by the Europeans who first came to Tibet because that name was used to refer to nearly all large dog breeds in the West. Early Western visitors to Tibet misnamed several of its breeds, such as the Tibetan terrier, which is not a terrier, and the Tibetan spaniel, which is not a spaniel. A better name for the breed might be the Tibetan mountain dog or—to encompass the landrace breed throughout its range—the Himalayan mountain dog.[2]

Description

[edit]

Appearance

[edit]
This side profile of Tü-Bo underscores the soundness and type that made him a sought-after stud dog.

It typically retains the hardiness which would be required for it to survive in Tibet, Ladakh and other high-altitude Himalayan regions.

Instinctive behaviors, including canine pack behavior, contributed to the survival of the breed in harsh environments. It is one of the few primitive dog breeds that retains a single estrus per year instead of two, even at much lower altitudes and in much more temperate climates than its native climate. This characteristic is also found in wild canids such as the wolf and other wild animals. Since its estrus usually takes place during late autumn, most Tibetan Mastiff puppies are born between December and January.[3]

Its double coat is long, subject to climate, and found in a wide variety of colors, including solid black, black and tan, various shades of red (from pale gold to deep red) and bluish-gray (dilute black), often with white markings. As of 2014, some breeders had begun marketing white Tibetan mastiffs.

The coat of a Tibetan Mastiff lacks the unpleasant big-dog smell that affects many large breeds. The coat, whatever its length or color(s), should shed dirt and odors. Although the dogs shed somewhat throughout the year, there is generally one great molt in late winter or early spring and sometimes another, lesser molt in the late summer or early autumn. (Sterilization of the dog may dramatically affect the coat as to texture, density and shedding pattern.)[4]

Tibetan Mastiffs are shown under one standard in the West, but separated by the Indian breed standard into two varieties: Lion Head (smaller; exceptionally long hair from forehead to withers, creating a ruff or mane) and Tiger Head (larger; shorter hair).

Temperament

[edit]

As a flock guardian dog in Tibet, and in the West, it uses all the usual livestock guardian tactics (e.g., barking, scent-marking perimeters) to warn away predators and avoid direct confrontations.[2]

As a socialized, more domestic dog, it can thrive in a spacious, fenced yard with a canine companion, but it is not an appropriate dog for apartment living. The Western-bred dogs are generally more easy-going, although still somewhat aloof with strangers. Through hundreds of years of selective breeding for a protective flock and family guardian, the breed has been prized for being a nocturnal sentry, keeping would-be predators and intruders at bay, and barking at unidentified sounds throughout the night. Leaving a Tibetan Mastiff outside all night with neighbors nearby is not recommended. They often sleep during the day, making them more active, alert, and aware at night.[2]

Like all flock guardian breeds, they are intelligent and stubborn to a fault, so obedience training is recommended (although it is only mildly successful with some individuals) since this is a strong-willed, powerful-bodied breed. Unless they are to be used exclusively as livestock guardians, socialization training is also critical with this breed, because of their reserved nature with strangers and guardian instincts. They can be excellent family dogs, depending on the family; owners must understand canine psychology and be able and willing to spend a lot of time and devotion to training their dogs. Lack of consistency can result in the creation of unpredictable dogs. The protectiveness of Tibetan Mastiffs requires alertness and planning by the owner, in order to avoid mishaps when the dog is merely reacting as a guardian. The breed is not recommended for novice dog owners.[2][5]

Health

[edit]

Many breeders claim a life expectancy of 10–16 years, but these claims are unsubstantiated. Some lines do produce long-lived dogs. Other, more closely inbred lines, produce short-lived, unhealthy dogs. The breed has fewer genetic health problems than many breeds, but cases can be found of hypothyroidism, entropion, ectropion, distichiasis, skin problems including allergies, autoimmune problems including demodex, Addison's disease, Cushing's disease, missing teeth, malocclusion (overbite, underbite, dry mouth), cardiac problems, seizures, epilepsy, progressive retinal atrophy (PRA), cataracts, and small ear canals with a tendency for infection. As with most large breeds, some will suffer with elbow or hip dysplasia.

Canine inherited demyelinative neuropathy (CIDN), an inherited condition, appeared in one of the prominent lines of Tibetan Mastiffs in the early 1980s.[6][full citation needed] Unfortunately, known carriers were bred extensively and are behind many lines still being actively bred.[citation needed] Because the mode of inheritance appears to be as a simple recessive, continued inbreeding can still produce affected puppies.

Hypothyroidism is fairly common in Tibetan Mastiffs, as it is in many large "northern" breeds. They should be tested periodically throughout their lives using a complete thyroid "panel". However, because the standard thyroid levels were established using domestic dog breeds, test results must be considered in the context of what is "normal" for the breed, not what is normal across all breeds. Many dogs of this breed will have "low" thyroid values, but no clinical symptoms. Vets and owners differ on the relative merits of medicating dogs which test "low", but are completely asymptomatic.

History

[edit]
Tibetan dog from the 1850s
Realistic representation/correction of Tibetan dog in 1904 "The German dogs". Heliogravure, Richard Strbel

Originally these dogs were used to protect Buddhist monasteries and monks of Tibet from animals such as bears, wolves and snow leopards.[7][2][5]

The Tibetan Mastiff is a phenotypically distinct dog breed that was bred as a flock guardian in the high altitudes of the Himalayas and the Tibetan Plateaus.[8][9]

Meer Izzut-oollah (1872) wrote of the Tibetan Mastiff:

The dogs of Tibet are twice the size of those seen in India, with large heads and hairy bodies. They are powerful animals...During the day they are kept chained up, and are let loose at night to guard their masters' house.[10]

In the early 20th century, the Prince of Wales, the future George V, introduced a pair of Tibetan Mastiffs to the United Kingdom. By 1906, the breed was prevalent enough in England to be shown at the Crystal Palace show. However, during the war years, the breed lost favor and focus and nearly died out in England.

The breed has been gaining in popularity worldwide since 1980. Although the breed is still considered somewhat uncommon, as more active breeders arose and produced adequate numbers of dogs, various registries and show organizations (FCI, AKC) began to recognize the breed. In 2008, the Tibetan Mastiff competed for the first time in the Westminster Kennel Club Dog Show.

Since AKC recognition, the number of active breeders has skyrocketed, leading to over-breeding of puppies, many of which are highly inbred and of questionable quality. Initially, the breed suffered because of the limited gene pool from the original stock.[citation needed] By 2015, due to excessive breeding and unsuitability of the breed as a pet in urban situations, prices in China for the best dogs had fallen to about $2,000, and both lower quality and crossbreed dogs were being abandoned.[11][5]

In 2011, a DNA study concluded that there was a genetic relationship between the Tibetan mastiff and the Great Pyrenees, Bernese Mountain Dog, Rottweiler and Saint Bernard, and that these large breed dogs are probably partially descended from the Tibetan Mastiff.[12] In 2014, a study added the Leonberger to the list of possible relatives.

Admixture with an unknown wolf-like canid

[edit]

The Tibetan Mastiff was able to adapt to the extreme highland conditions of the Tibetan Plateau very quickly compared with other mammals such as the yak, Tibetan antelope, snow leopard, and the wild boar. The Tibetan mastiff's ability to avoid hypoxia in high altitudes, due to its higher hemoglobin levels compared with low-altitude dogs, was due to prehistoric interbreeding.[13][14] In 2020, a genomic analysis indicates that a ghost population of an unknown wolf-like canid which is deeply-diverged from modern Holarctic wolves and dogs has contributed the EPAS1 allele found in both Himalayan wolves and dogs, and this allows them to live in high altitudes.[15]

[edit]
[edit]
  • A Tibetan Mastiff named "Max" is the central antagonist in the 1993 horror film Man's Best Friend. At least five different dogs were used in filming.
  • Ulysses, one of the main characters in the 2008 Christian animated film At Jesus' Side, is a Tibetan Mastiff.
  • A Tibetan Mastiff is the subject of the 2011 animated film The Tibetan Dog.
  • Mouse, a "Tibetan Temple dog" (a semi-divine creature that closely resembles a Mastiff), is the canine companion of the titular character of the Dresden Files book series.
  • The animated film Rock Dog featured two Tibetan Mastiffs named "Bodi" and "Khampa" (voiced by Luke Wilson and J. K. Simmons, respectively).
  • In the 2018 animated television series, Craig of the Creek, the character Wildernessa rides around on a Tibetan Mastiff named Cheesesticks.
  • In the 2011 film Old Dog by director Pema Tseden, the Tibetan Mastiff of a herder family is coveted by several characters to be traded to rich Han Chinese in the eastern part of the country.
  • The 2019 video game Indivisible features Lanshi, a Tibetan Mastiff, as a playable character.
  • The 2019 manga Ginga Densetsu Noah features three Tibetan Mastiffs as the main antagonists.
  • The 2020 video game Ghost of Tsushima, Tibetan Mastiff is also known Bankhar Dog, were introduced as guard dog of Mongol army.
  • The Pokémon character Mabosstiff, first featured in Scarlet and Violet versions, is heavily based on a Tibetan Mastiff.

See also

[edit]

Notes

[edit]
  1. ^ Tibetan: འབྲོག་ཁྱི། འདོགས་ཁྱི། སེང་ཁྱི།, romanized: Drok chi/kyi; 'Dok kyi; Seng kyi, lit.'Nomad-dog; chaining dog; lion-dog', THL: Do khyi, Wylie: 'brog khyi; 'dogs khyi; seng khyi

References

[edit]
  1. ^ a b FCI breed standard
  2. ^ a b c d e Messerchmidt, Don (2010). Discovering the Big Dogs of Tibet and the Himalayas: A personal journey. [page needed]
  3. ^ "Tibetan mastiff". American Kennel Club. Dog breed information. Retrieved 9 March 2019.
  4. ^ citation needed
  5. ^ a b c Tibetan Dogs: A complete anthology of the breeds. Vintage Dog Books. 18 November 2010. ISBN 978-1-4455-2671-3. [page needed]
  6. ^ "The Tibetan Mastiff" by Ann Rohrer and Cathy J. Flamholtz
  7. ^ Palika, Liz (2007). The Howell Book of Dogs: The definitive reference to 300 breeds and varieties. John Wiley & Sons. p. 374. ISBN 978-0-470-17585-9 – via archive.org.
  8. ^ Messerschmidt, D.M.R. (1983). "The Tibetan Mastiff: Canine sentinels of the range". Rangelands. Vol. 5. pp. 172–174.
  9. ^ Li, Q.; Liu, Z.; Li, Y.; Zhao, X.; Dong, L.; Pan, Z.; Sun, Y.; Li, N.; Xu, Y.; Xie, Z. (2008). "Origin and phylogenetic analysis of Tibetan mastiff based on the mitochondrial DNA sequence". Journal of Genetics and Genomics. 35 (6): 335–340. doi:10.1016/S1673-8527(08)60049-1. PMID 18571121.
  10. ^ Izzut-oollah, Meer (1872). Travels in Central Asia in the Years 1812–13. Translated by Henderson. Calcutta, IN: Foreign Department Press. p. 15.
  11. ^ "Once-prized Tibetan mastiffs are discarded as fad ends in China". The New York Times. World/Asia. New York, NY. 2015-04-18. Retrieved 27 February 2017.
  12. ^ Li, Y.; Zhao, X.; Pan, Z.; Xie, Z.; Liu, H.; Xu, Y.; Li, Q. (2011). "The origin of the Tibetan mastiff and species identification of Canis based on mitochondrial cytochrome c oxidase subunit I (COI) gene and COI barcoding". Animal. 5 (12): 1868–73. Bibcode:2011Anim....5.1868L. doi:10.1017/S1751731111001042. PMID 22440462.
  13. ^ Miao, Benpeng; Wang, Zhen; Li, Yixue (2016). "Genomic analysis reveals hypoxia adaptation in the Tibetan mastiff by introgression of the grey wolf from the Tibetan plateau". Molecular Biology and Evolution. 34 (3): 734–743. doi:10.1093/molbev/msw274. PMID 27927792. S2CID 47507546.
  14. ^ Signore, Anthony V.; Yang, Ying-Zhong; Yang, Quan-Yu; Qin, Ga; Moriyama, Hideaki; Ge, Ri-Li; Storz, Jay F. (2019). "Adaptive changes in hemoglobin function in high-altitude Tibetan canids were derived via gene conversion and introgression". Molecular Biology and Evolution. 36 (10): 2227–2237. doi:10.1093/molbev/msz097. PMC 6759075. PMID 31362306.
  15. ^ Wang, Ming-Shan; Wang, Sheng; Li, Yan; Jhala, Yadvendradev; Thakur, Mukesh; Otecko, Newton O.; Si, Jing-Fang; Chen, Hong-Man; Shapiro, Beth; Nielsen, Rasmus; Zhang, Ya-Ping; Wu, Dong-Dong (2020). "Ancient Hybridization with an Unknown Population Facilitated High-Altitude Adaptation of Canids". Molecular Biology and Evolution. 37 (9): 2616–2629. doi:10.1093/molbev/msaa113. PMID 32384152.
[edit]